Gene regulation of CAM & C4 photosynthesis Single-nuclei sequencing of the regulatory architecture behind integrated C4+CAM photosynthesis in Portulaca. Pelargonium CAM genome project Investigating the genetic basis of CAM in southern African Pelargonium species. Phylogenomics of Portullugo & beyond Methods for reconstructing evolutionary histories from phylogenomic and museum-specimen data. Comparative anatomy, physiology & machine learning Linking photosynthetic anatomy, physiology, and gene expression across vascular plants.